Touchpad natural scroll
I can't find a way to enable natural scroll with the touchpad - down movement on the touchpad means the screen moves down - in Firefox. I've gone into about:config and set mousewheel.default.delta_multiplier_y to -100, but that only affects the mouse, not the trackpad.
Solution choisie
I did have global setting for touchpad scroll directions enabled, but firefox wasn't agreeing with them - I refreshed firefox and then the browser decided to be more cooperative.
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Windows 10 has global touchpad and mouse settings that enable natural scrolling. If you are using a public laptop that prohibits changing these settings, then you could try and boot Firefox on a live OS.
Solution choisie
I did have global setting for touchpad scroll directions enabled, but firefox wasn't agreeing with them - I refreshed firefox and then the browser decided to be more cooperative.